For outstanding scientific leadership in nuclear physics and foundational work in developing and applying nuclear density functional theory to atomic nuclei
For pioneering research in ecosystem theory, ecological modeling, error analysis, hierarchy theory, and landscape ecology and for the development of basic applications in risk assessment and regional environmental analysis.
For original studies of the genetic effects of radiation in mammals. A world authority on mammalian mutagenesis, he and co-workers provided the experimental basis for estimating the genetic hazards of radiation to man and for the corresponding recommendations of national and international standards bodies